Publications

Pontzer, H., Rimbach, R., Paltan, J., Ivory, E., Kendall, C. “Air temperature and diet influence body composition and water turnover in zoo-living African elephants (Loxodonta africana)“. The Royal Society Open Sci. (October 2020). Full Text

Bethancourt, H.J., Swanson, S.Z., Nzunza, R., Huanca, T., Conde, E., Kenney, W.L., Young, S.L., Ndiema, E., Braun, D., Pontzer, H., Rosinger1, A.Y. “Hydration in relation to water insecurity, heat index, and lactation status in two small-scale populations in hot-humid and hot-arid environments“. American Journal of Human Biology (June 2020): Full Text

Wood, B.M., Pontzer, H., Harris, J.A., Mabulla, A.Z.P., … Raichlen, D.A. “Step counts from satellites: methods for integrating accelerometer and GPS data for more accurate measures of pedestrian travel”. Journal of the Measurement of Physical Behaviour 3 (March 2020): 58-66. Full Text

Stoner, L., Pontzer, H., Barone Gibbs, B., Moore, J.B., Castro, N., Skidmore, S., Williams, M.A., Hamlin, M.J., Faulkner, J. “Fitness and fatness are both associated with cardiometabolic risk in pre-adolescents“. Journal of Pediatrics 217 (February 2020): 39–45. Full Text

Hora, M. Pontzer, H., Wall-Scheffler, C.M., Sládek, V. “Dehydration and persistence hunting in Homo erectus“. Journal of Human Evolution 138 (January 2020): 102682. Full Text

Urlacher, S.S., Snodgrass, J.J., Dugas, L.R., Sugiyama, L.S., Liebert, M.A., Joyce, C.J., Pontzer, H. “Constraint and tradeoffs regulate energy expenditure during childhood“. Science Advances 5 (December 2019): eaax1065. Full Text

Swanson, Z., Pontzer, H. “Water Turnover Among Human Populations: Effects of Environment and Lifestyle”. American Journal of Human Biology 32 (November 2019), e23365. Full Text

Speakman, J.R., Pontzer, H., Rood, J., Sagayama, H., Schoeller, D.A., Westerterp, K.R., Wong, W.W., Yamada, Y., Loech, C., Alford, A. “The IAEA international doublylabelled water database: aims, scope and procedures“. Annals of Nutrition and Metabolism 75 (November 2019): 114-118. Full Text

Horiuchi, M., Handa-Kirihara, Y., Fukuoka, Y., Pontzer, H. “Sex differences in respiratory and circulatory cost and arterial oxygen saturation during hypoxic walking”. Scientific Reports 9 (July 2019): 9550. Full Text

Pontzer, H. “Step on it”. New Scientist. (June 2019): 34-37. Full Text

Thurber, C., Dugas, L.R., Ocobock, C., Carlson, B., Speakman, J.R., Pontzer, H. “Extreme events reveal an  alimentary limit on sustained maximal human energy expenditure”. Science Advances 5 (June 2019), eaaw0341. Full Text

Rosinger, A.Y., Pontzer, H., Raichlen, D.A., … Sands, J.M. “Human age-related decline in urine concentrating ability may not be universal: Hydration biomarkers from the US and two small-scale societies”. American Journal of Physical Anthropology 168, (April 2019): 705-716. Full Text

Gurven, M., Trumble, B., Stieglitz, J., … Pontzer, H. “Methodological differences cannot explain associations between health, anthropometrics, and excess resting metabolic rate”. American Journal of Physical Anthropology 169 (March 2019): 197-198. Full Text

Christopher, L., Madimenos, F., Bribiescas, R., Urlacher, S., Snodrass, J.J., Sugiyama, L., Pontzer, H. “High energy requirements and water throughput of adult Shuar foragerhorticulturalists of Amazonian Ecuador”. American Journal of Human Biology 31 (February 2019), e23223. Full Text

Ocobock, C., Overbeck, A., Carlson, C., Royer, C., Mervenne, A., *Thurber, C., … Pontzer, H. “Sustained high levels of physical activity lead to improved performance among “Race Across the USA” athletes”. American Journal of Physical Anthropology 168 (January 2019): 789-794. Full Text

Pontzer, H. “Evolved to exercise”. Scientific American. (January 2019): 22-29.

Pontzer, H. “Method and rationale for recalculating dilution spaces to a single, common time point in doubly labeled water studies”. European Journal of Clinical Nutrition. Volume 72, (December 2018): 1620-1624.

Pontzer, H., Raichlen, D.A., Wood, B.M. “Hunter-gatherers as models in public health”. Obesity Reviews 19 (December 2018) (Suppl 1): 24-35. Full Text 

Pontzer, H. “Energy Constraint as a Novel Mechanism Linking Exercise and Health.” Physiology (Bethesda, Md.) 33.6 (November 2018): 384-393. Full Text

Pontzer, H. “Alternative Energy Physiological evolution in the human lineage.” Natural History 126.8 (September 1, 2018): 37-39. (Academic Article)

Gagnon, CM, Steiper, ME, and Pontzer, H. “Elite swimmers do not exhibit a body mass index trade-off across a wide range of event distances.” Proceedings. Biological Sciences 285.1882 (July 4, 2018). Full Text

Finestone, EM, Brown, MH, Ross, SR, and Pontzer, H. “Great ape walking kinematics: Implications for hominoid evolution.” American Journal of Physical Anthropology 166.1 (May 2018): 43-55. Full Text

Urlacher, SS, Ellison, PT, Sugiyama, LS, Pontzer, H, Eick, G, Liebert, MA, Cepon-Robins, TJ, Gildner, TE, and Snodgrass, JJ. “Tradeoffs between immune function and childhood growth among Amazonian forager-horticulturalists.” Proceedings of the National Academy of Sciences of the United States of America 115.17 (April 9, 2018): E3914-E3921. Full Text

Kozma, EE, Webb, NM, Harcourt-Smith, WEH, Raichlen, DA, D’Août, K, Brown, MH, Finestone, EM, Ross, SR, Aerts, P, and Pontzer, H. “Hip extensor mechanics and the evolution of walking and climbing capabilities in humans, apes, and fossil hominins.” Proceedings of the National Academy of Sciences of the United States of America 115.16 (April 2, 2018): 4134-4139. Full Text

Pontzer, H, Raichlen, DA, Basdeo, T, Harris, JA, Mabulla, AZP, and Wood, BM. “Mechanics of archery among Hadza hunter-gatherers.” Journal of Archaeological Science: Reports 16 (December 2017): 57-64. Full Text

Horiuchi, M, Fukuoka, Y, Handa, Y, Abe, D, and Pontzer, H. “Measuring the Energy of Ventilation and Circulation during Human Walking using Induced Hypoxia.” Scientific Reports 7.1 (July 10, 2017): 4938-null. Full Text

Pontzer, H. “Economy and Endurance in Human Evolution.” Current Biology : Cb 27.12 (June 2017): R613-R621. (Review) Full Text

Edwards, W, Lonsdorf, EV, and Pontzer, H. “Total energy expenditure in captive capuchins (Sapajus apella).” American Journal of Primatology 79.5 (May 2017). Full Text

Raichlen, DA, Pontzer, H, Harris, JA, Mabulla, AZP, Marlowe, FW, Josh Snodgrass, J, Eick, G, Colette Berbesque, J, Sancilio, A, and Wood, BM. “Physical activity patterns and biomarkers of cardiovascular disease risk in hunter-gatherers.” American Journal of Human Biology : the Official Journal of the Human Biology Council 29.2 (March 2017). Full Text

Pontzer, H. “The crown joules: energetics, ecology, and evolution in humans and other primates.” Evolutionary Anthropology 26.1 (January 2017): 12-24. Full Text

Sparrow, LM, Pellatt, E, Yu, SS, Raichlen, DA, Pontzer, H, and Rolian, C. “Gait changes in a line of mice artificially selected for longer limbs.” Peerj 5 (January 2017): e3008-null. Full Text

Hora, M, Soumar, L, Pontzer, H, and Sládek, V. “Body size and lower limb posture during walking in humans.” Plos One 12.2 (January 2017): e0172112-null. Full Text

Pontzer, H. “The Exercise Paradox.” Scientific American 316.2 (January 2017): 26-31. Full Text

Gurven, MD, Trumble, BC, Stieglitz, J, Yetish, G, Cummings, D, Blackwell, AD, Beheim, B, Kaplan, HS, and Pontzer, H. “High resting metabolic rate among Amazonian forager-horticulturalists experiencing high pathogen burden.” American Journal of Physical Anthropology 161.3 (November 2016): 414-425. Full Text

Pontzer, H. “Behavior: Knowing When to Walk Away, Knowing When to Run.” Current Biology : Cb 26.15 (August 2016): R717-R718. Full Text

Pontzer, H, Brown, MH, Raichlen, DA, Dunsworth, H, Hare, B, Walker, K, Luke, A, Dugas, LR, Durazo-Arvizu, R, Schoeller, D, Plange-Rhule, J, Bovet, P, Forrester, TE, Lambert, EV, Thompson, ME, Shumaker, RW, and Ross, SR. “Metabolic acceleration and the evolution of human brain size and life history.” Nature 533.7603 (May 4, 2016): 390-392. Full Text

Yetish, G, Kaplan, H, Gurven, M, Wood, B, Pontzer, H, Manger, PR, Wilson, C, McGregor, R, and Siegel, JM. “Response to de la Iglesia et al.” Current Biology : Cb 26.7 (April 2016): R273-R274. (Letter) Full Text

Laird, MF, Vogel, ER, and Pontzer, H. “Chewing efficiency and occlusal functional morphology in modern humans.” Journal of Human Evolution 93 (April 2016): 1-11. Full Text

Pontzer, H, Durazo-Arvizu, R, Dugas, LR, Plange-Rhule, J, Bovet, P, Forrester, TE, Lambert, EV, Cooper, RS, Schoeller, DA, and Luke, A. “Constrained Total Energy Expenditure and Metabolic Adaptation to Physical Activity in Adult Humans.” Current Biology : Cb 26.3 (February 2016): 410-417. Full Text

Pontzer, H. “A unified theory for the energy cost of legged locomotion.” Biology Letters 12.2 (February 2016): 20150935-null. Full Text

Yetish, G, Kaplan, H, Gurven, M, Wood, B, Pontzer, H, Manger, PR, Wilson, C, McGregor, R, and Siegel, JM. “Natural sleep and its seasonal variations in three pre-industrial societies.” Current Biology : Cb 25.21 (November 2015): 2862-2868. Full Text

Pontzer, H. “Energy Expenditure in Humans and Other Primates: A New Synthesis.” Annual Review of Anthropology 44.1 (October 21, 2015): 169-187. Full Text

Pontzer, H, Raichlen, DA, Wood, BM, Emery Thompson, M, Racette, SB, Mabulla, AZP, and Marlowe, FW. “Energy expenditure and activity among Hadza hunter-gatherers.” American Journal of Human Biology : the Official Journal of the Human Biology Council 27.5 (September 2015): 628-637. Full Text

Pontzer, H. “Constrained Total Energy Expenditure and the Evolutionary Biology of Energy Balance.” Exercise and Sport Sciences Reviews 43.3 (July 2015): 110-116. (Review) Full Text

Warrener, AG, Lewton, KL, Pontzer, H, and Lieberman, DE. “A wider pelvis does not increase locomotor cost in humans, with implications for the evolution of childbirth.” Plos One 10.3 (January 2015): e0118903-null. Full Text

Wood, BM, Pontzer, H, Raichlen, DA, and Marlowe, FW. “Mutualism and manipulation in Hadza–honeyguide interactions.” Evolution and Human Behavior 35.6 (November 2014): 540-546. Full Text

Pontzer, H, Suchman, K, Raichlen, DA, Wood, BM, Mabulla, AZP, and Marlowe, FW. “Foot strike patterns and hind limb joint angles during running in Hadza hunter-gatherers.” Journal of Sport and Health Science 3.2 (June 2014): 95-101. Full Text

Raubenheimer, D, Rothman, JM, Pontzer, H, and Simpson, SJ. “Macronutrient contributions of insects to the diets of hunter-gatherers: a geometric analysis.” Journal of Human Evolution 71 (June 2014): 70-76. Full Text

Pontzer, H, Raichlen, DA, Gordon, AD, Schroepfer-Walker, KK, Hare, B, O’Neill, MC, Muldoon, KM, Dunsworth, HM, Wood, BM, Isler, K, Burkart, J, Irwin, M, Shumaker, RW, Lonsdorf, EV, and Ross, SR. “Primate energy expenditure and life history.” Proceedings of the National Academy of Sciences of the United States of America 111.4 (January 13, 2014): 1433-1437. Full Text

Pontzer, H. “Early Hominin Paleoecology.” American Journal of Human Biology 26.1 (January 1, 2014): 103-103. (Review) Full Text

Pontzer, H, Raichlen, DA, and Rodman, PS. “Bipedal and quadrupedal locomotion in chimpanzees.” Journal of Human Evolution 66 (January 2014): 64-82. Full Text

Raichlen, DA, Wood, BM, Gordon, AD, Mabulla, AZP, Marlowe, FW, and Pontzer, H. “Evidence of Levy walk foraging patterns in human hunter-gatherers.” Proceedings of the National Academy of Sciences of the United States of America 111.2 (January 2014): 728-733. Full Text

Foster, AD, Raichlen, DA, and Pontzer, H. “Muscle force production during bent-knee, bent-hip walking in humans.” Journal of Human Evolution 65.3 (September 2013): 294-302. Full Text

Barak, MM, Lieberman, DE, Raichlen, D, Pontzer, H, Warrener, AG, and Hublin, J-J. “Trabecular evidence for a human-like gait in Australopithecus africanus.” Plos One 8.11 (January 2013): e77687-null. Full Text

Raichlen, DA, Pontzer, H, and Shapiro, LJ. “A new look at the Dynamic Similarity Hypothesis: the importance of swing phase.” Biology Open 2.10 (January 2013): 1032-1036. Full Text

Pontzer, H. “Ecological Energetics in Early Homo.” Current Anthropology 53.S6 (December 2012): S346-S358. Full Text

Dunsworth, HM, Warrener, AG, Deacon, T, Ellison, PT, and Pontzer, H. “Metabolic hypothesis for human altriciality.” Proceedings of the National Academy of Sciences of the United States of America 109.38 (September 2012): 15212-15216. Full Text

Pontzer, H. “Relating ranging ecology, limb length, and locomotor economy in terrestrial animals.” Journal of Theoretical Biology 296 (March 2012): 6-12. Full Text

Pontzer, H, Raichlen, DA, Wood, BM, Mabulla, AZP, Racette, SB, and Marlowe, FW. “Hunter-gatherer energetics and human obesity.” Plos One 7.7 (January 2012): e40503-null. Full Text

Pontzer, H, Scott, JR, Lordkipanidze, D, and Ungar, PS. “Dental microwear texture analysis and diet in the Dmanisi hominins.” Journal of Human Evolution 61.6 (December 2011): 683-687. Full Text

Orkin, JD, and Pontzer, H. “The Narrow Niche hypothesis: gray squirrels shed new light on primate origins.” American Journal of Physical Anthropology 144.4 (April 2011): 617-624. Full Text

Cowgill, LW, Warrener, A, Pontzer, H, and Ocobock, C. “Waddling and toddling: the biomechanical effects of an immature gait.” American Journal of Physical Anthropology 143.1 (September 2010): 52-61. Full Text

Pontzer, H, Raichlen, DA, Shumaker, RW, Ocobock, C, and Wich, SA. “Metabolic adaptation for low energy throughput in orangutans.” Proceedings of the National Academy of Sciences of the United States of America 107.32 (August 2, 2010): 14048-14052. Full Text

Pontzer, H, Rolian, C, Rightmire, GP, Jashashvili, T, Ponce de León, MS, Lordkipanidze, D, and Zollikofer, CPE. “Locomotor anatomy and biomechanics of the Dmanisi hominins.” Journal of Human Evolution 58.6 (June 2010): 492-504. Full Text

Raichlen, DA, Pontzer, H, Shapiro, LJ, and Sockol, MD. “Understanding hind limb weight support in chimpanzees with implications for the evolution of primate locomotion.” American Journal of Physical Anthropology 138.4 (April 2009): 395-402. Full Text

Pontzer, H, Holloway IV, JH, Raichlen, DA, and Lieberman, DE. “Control and function of arm swing in human walking and running (Journal of Experimental Biology 212 (523-534)).” Journal of Experimental Biology 212.6 (March 15, 2009): 894-null. Full Text

Pontzer, H, Holloway, JH, Raichlen, DA, and Lieberman, DE. “Control and function of arm swing in human walking and running.” The Journal of Experimental Biology 212.Pt 4 (February 2009): 523-534. Full Text

Pontzer, H, Allen, V, and Hutchinson, JR. “Biomechanics of running indicates endothermy in bipedal dinosaurs.” Plos One 4.11 (January 2009): e7783-null. Full Text

Pontzer, H, Raichlen, DA, and Sockol, MD. “The metabolic cost of walking in humans, chimpanzees, and early hominins.” Journal of Human Evolution 56.1 (January 2009): 43-54. Full Text

Pontzer, H, and Kamilar, JM. “Great ranging associated with greater reproductive investment in mammals.” Proceedings of the National Academy of Sciences of the United States of America 106.1 (January 2009): 192-196. Full Text

Carter, ML, Pontzer, H, Wrangham, RW, and Peterhans, JK. “Skeletal pathology in Pan troglodytes schweinfurthii in Kibale National Park, Uganda.” American Journal of Physical Anthropology 135.4 (April 2008): 389-403. Full Text

Raichlen, DA, Pontzer, H, and Sockol, MD. “The Laetoli footprints and early hominin locomotor kinematics.” Journal of Human Evolution 54.1 (January 2008): 112-117. Full Text

Lordkipanidze, D, Jashashvili, T, Vekua, A, Ponce de León, MS, Zollikofer, CPE, Rightmire, GP, Pontzer, H, Ferring, R, Oms, O, Tappen, M, Bukhsianidze, M, Agusti, J, Kahlke, R, Kiladze, G, Martinez-Navarro, B, Mouskhelishvili, A, Nioradze, M, and Rook, L. “Postcranial evidence from early Homo from Dmanisi, Georgia.” Nature 449.7160 (September 2007): 305-310. Full Text

Sockol, MD, Raichlen, DA, and Pontzer, H. “Chimpanzee locomotor energetics and the origin of human bipedalism.” Proceedings of the National Academy of Sciences of the United States of America 104.30 (July 16, 2007): 12265-12269. Full Text

Pontzer, H. “Effective limb length and the scaling of locomotor cost in terrestrial animals.” The Journal of Experimental Biology 210.Pt 10 (May 2007): 1752-1761. Full Text

Pontzer, H. “Predicting the energy cost of terrestrial locomotion: a test of the LiMb model in humans and quadrupeds.” The Journal of Experimental Biology 210.Pt 3 (February 2007): 484-494. Full Text

Lieberman, DE, Raichlen, DA, Pontzer, H, Bramble, DM, and Cutright-Smith, E. “The human gluteus maximus and its role in running.” The Journal of Experimental Biology 209.Pt 11 (June 2006): 2143-2155. Full Text

Pontzer, H, and Wrangham, RW. “Ontogeny of Ranging in Wild Chimpanzees.” International Journal of Primatology 27.1 (February 2006): 295-309. Full Text

Pontzer, H, Lieberman, DE, Momin, E, Devlin, MJ, Polk, JD, Hallgrímsson, B, and Cooper, DM. “Trabecular bone in the bird knee responds with high sensitivity to changes in load orientation.” The Journal of Experimental Biology 209.Pt 1 (January 2006): 57-65. Full Text

Pontzer, H. “A new model predicting locomotor cost from limb length via force production.” The Journal of Experimental Biology 208.Pt 8 (April 2005): 1513-1524. Full Text

Pontzer, H, and Wrangham, RW. “Climbing and the daily energy cost of locomotion in wild chimpanzees: implications for hominoid locomotor evolution.” Journal of Human Evolution 46.3 (March 2004): 317-335. Full Text