Broadly, my research addresses the physiological, molecular and genetic mechanisms of plant adaptive responses to stressful environments. A grand challenge of Evolutionary Biology is to understand how organisms mitigate environmental perturbations. Increased prevalence and magnitude of environmental stress in the face of global climate change provides a strong impetus for understanding plant stress responses. Using the model plant Boechera stricta, I combine quantitative genetics, population genomics, physiology and ecological genetics to understand how plants have evolved in response to extreme environments. Through this work I aim to: 1) identify the polymorphic genes/pathways/networks that modulate adaptive phenotypic states; and 2) elucidate the genetic architecture of fitness components; and 3) understand the past, present and future evolutionary trajectories of plant responses to stressful environmental shifts.